Joseph Tomkins

Dr, BSc Sheff., PhD Liv.

  • The University of Western Australia (M092), 35 Stirling Highway, Room 2.15, Zoology Building, Perth campus

    6009 Perth


  • 2779 Citations
  • 30 h-Index
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Personal profile


I have an Honours degree in Zoology from the University of Sheffield, and a PhD from the University of Liverpool. I have had Research Fellowships in Australia and the UK. In the UK I spent 2000-2005 at the University of St Andrews and one year at the University of Exeter. At the University of Western Australia I was employed on fellowships from 1997 - 2000 and again from 2005 to 2011. I was an ARC Future Fellow from 2011-2015.

Roles and responsibilities

Research Committee Member.

Future research

My Future Fellowship Research was on the evolution of threshold traits and covered traits as diverse as mate-choice and polyandry to the spread of disease by ticks. This research is ongoing.

I have received ARC funding (starting 2017) to develop an empirical understanding of anisogamy. I am intrested in a breadth of subjects mostly centred around sexual selection, mutations and threshold evolution.

Teaching overview

Honours supervision

3rd Year Behavioural Ecology

3rd Year Evolutionary Processes

1st Year 'The Darwinian Revolution'


My research is in sexual selection. I am particularly interested in the genetic architecture of sexually selected and life-history traits, allometry, phenotypic plasticity and developmental integration. I have a broad interest in threshold traits and their evolutionary quirks. I am currently developing methodologies for experimentally manipulating anisogamy.


I speak English, sometimes in hilarious accents. I speak Franglais and Spanglish with equal ease.


  • Allometry
  • Alternative reproductive tactics
  • Alternative strategies
  • Genetic variation
  • Inbreeding
  • Sexual selection
  • Threshold traits

Fingerprint Dive into the research topics where Joseph Tomkins is active. These topic labels come from the works of this person. Together they form a unique fingerprint.

  • 2 Similar Profiles
beetle Earth & Environmental Sciences
sexual selection Agriculture & Biology
allometry Agriculture & Biology
Onthophagus taurus Agriculture & Biology
dimorphism Agriculture & Biology
evolutionarily stable strategy Earth & Environmental Sciences
sperm competition Earth & Environmental Sciences
dung beetles Agriculture & Biology

Network Recent external collaboration on country level. Dive into details by clicking on the dots.

Research Output 1999 2019

Evaluating the genetic architecture of quantitative traits via selection followed by inbreeding

Dugand, R. J., Kennington, W. J. & Tomkins, J. L., 9 Apr 2019, In : Heredity. 12 p.

Research output: Contribution to journalArticle

Body Size
Drosophila melanogaster
Gene Frequency

It's not all black and white: Investigating colour polymorphism in manta rays across Indo-Pacific populations

Venables, S. K., Marshall, A. D., Germanov, E. S., Perryman, R. J. Y., Tapilatu, R. F., Gede Hendrawan, I., Flam, A. L., Van Keulen, M., Tomkins, J. L. & Kennington, W. J., 1 Jan 2019, In : Proceedings of the Royal Society B: Biological Sciences. 286, 1912, 20191879.

Research output: Contribution to journalArticle


Male responses to sperm competition when rivals vary in number and familiarity

Lymbery, S. J., Tomkins, J. L. & Simmons, L. W., 30 Jan 2019, In : Proceedings of the Royal Society B: Biological Sciences. 286, 1895, 20182589.

Research output: Contribution to journalArticle

sperm competition
kin recognition

Male semelparity and multiple paternity confirmed in an arid-zone dasyurid

Hayes, G. L. T., Simmons, L. W., Dugand, R. J., Mills, H. R., Roberts, J. D., Tomkins, J. L. & Fisher, D. O., 21 Apr 2019, In : Journal of Zoology.

Research output: Contribution to journalArticle

arid zones
litters (young animals)
3 Citations (Scopus)

Molecular evidence supports a genic capture resolution of the lek paradox

Dugand, R., Tomkins, J. & Kennington, W., 25 Mar 2019, In : Nature Communications. 10, 1359.

Research output: Contribution to journalArticle

Open Access


Chlamydomonas flow data files

Seed, C. (Creator), Tomkins, J. (Creator), Dryad Digital Repository, 4 Oct 2016


Data from: Towards an evolutionary understanding of questing behaviour in the tick Ixodes ricinus

Tomkins, J. (Creator), Aungier, J. (Creator), Hazel, W. N. (Creator), Gilbert, L. (Creator), Dryad Digital Repository, 21 Oct 2014


Projects 2000 2019