Joseph Tomkins

Dr, BSc Sheff., PhD Liv.

  • The University of Western Australia (M092), 35 Stirling Highway, Room 2.15, Zoology Building, Perth campus

    6009 Perth


  • 2435 Citations
  • 27 h-Index
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Personal profile


I have an Honours degree in Zoology from the University of Sheffield, and a PhD from the University of Liverpool. I have had Research Fellowships in Australia and the UK. In the UK I spent 2000-2005 at the University of St Andrews and one year at the University of Exeter. At the University of Western Australia I was employed on fellowships from 1997 - 2000 and again from 2005 to 2011. I was an ARC Future Fellow from 2011-2015.

Roles and responsibilities

Research Committee Member.

Future research

My Future Fellowship Research was on the evolution of threshold traits and covered traits as diverse as mate-choice and polyandry to the spread of disease by ticks. This research is ongoing.

I have received ARC funding (starting 2017) to develop an empirical understanding of anisogamy. I am intrested in a breadth of subjects mostly centred around sexual selection, mutations and threshold evolution.

Teaching overview

Honours supervision

3rd Year Behavioural Ecology

3rd Year Evolutionary Processes

1st Year 'The Darwinian Revolution'


My research is in sexual selection. I am particularly interested in the genetic architecture of sexually selected and life-history traits, allometry, phenotypic plasticity and developmental integration. I have a broad interest in threshold traits and their evolutionary quirks. I am currently developing methodologies for experimentally manipulating anisogamy.


I speak English, sometimes in hilarious accents. I speak Franglais and Spanglish with equal ease.


  • Allometry
  • Alternative reproductive tactics
  • Alternative strategies
  • Genetic variation
  • Inbreeding
  • Sexual selection
  • Threshold traits

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Projects 2000 2019

Research Output 1999 2018

Evolutionary divergence in competitive mating success through female mating bias for good genes

Dugand, R. J., Jason Kennington, W. & Tomkins, J. L., 23 May 2018, In : Science Advances. 4, 5, Y.

Research output: Contribution to journalArticle

Open Access
Drosophila melanogaster

Fitness consequences of threshold trait expression subject to environmental cues

Michalczyk, Ł., Dudziak, M., Radwan, J. & Tomkins, J. L., 27 Jun 2018, In : Proceedings of the Royal Society B: Biological Sciences. 285, 1881, 20180783.

Research output: Contribution to journalArticle

environmental cue
evolutionarily stable strategy

Living with fire: Ecology and genetics of the dasyurid mammal Dasykaluta rosamondae

Hayes, G. L., 2018, (Unpublished)

Research output: ThesisDoctoral Thesis

fire ecology
arid zones
Chlamydomonas reinhardtii
vegetative cells
Germ Cells

State-dependent changes in risk-taking behaviour as a result of age and residual reproductive value

Moschilla, J. A., Tomkins, J. L. & Simmons, L. W., 1 Aug 2018, In : Animal Behaviour. 142, p. 95-100 6 p.

Research output: Contribution to journalArticle

risk behavior
residual effects
Teleogryllus oceanicus


Data from: Towards an evolutionary understanding of questing behaviour in the tick Ixodes ricinus

Tomkins, J. (Creator), Aungier, J. (Creator), Hazel, W. N. (Creator), Gilbert, L. (Creator), Dryad Digital Repository, 21 Oct 2014


Chlamydomonas flow data files

Seed, C. (Creator), Tomkins, J. (Creator), Dryad Digital Repository, 4 Oct 2016